Ori, the protagonist of the game, falls from the Spirit Tree and is adopted by Naru, who raises Ori as her own. When a disastrous event occurs causing the forest to wither and Naru to die, Ori is left to explore the forest. Ori eventually encounters Sein, who begins to guide Ori on an adventure to restore the forest through the recovery of the light of three main elements supporting the balance of the forest: waters, winds and warmth.